AO Univ. Prof. Hans-Christoph Winkler
Konrad-Lorenz-Institut für Vergleichende Verhaltensforschung
Department für Integrative Biologie und Evolution
Veterinärmedizinische Universität Wien
Savoyenstr. 1a
A-1160 Wien
T +43 (1) 489 09 15 822
F +43 (1) 489 09 15 801
E-Mail an Hans-Christoph Winkler senden [Link 1]
Research
- I am interested in a variety of topics integrating behaviour, ecology, and morphology, particularly of birds, with an emphasis on comparative approaches.
- I am especially paying attention to the influences of the environment on behaviour, especially communication, mating systems and cognition, and on the interdependences between morphology and behaviour.
Research Summary
The main goal of my research is to understand how ecology and behaviour interact on the mechanistic and evolutionary level. I am particularly fascinated by the question how morphology mediates these interactions, and I am equally captivated by the evolutionary and ecological processes that generate and maintain biological diversity. That diversity forms also the base for applying comparative methods to analyze the evolution of behavioural and morphological traits. My current work concerns the relationship between intra-specific and inter-specific variation in the course of diversification, the role of habitat in acoustical and optical communication, the evolution of mating systems and social interactions, and the ecological conditions for the evolution of cognitive skills, and how they in turn influence evolutionary processes. Finally, I am interested to apply scientific thinking and results to conservation.
Current Projects
Curriculum Vitae
1993 | Tit. ao. Prof., University of Vienna |
1980 | Habilitation, University of Vienna |
1971 | Dr. phil, University of Vienna |
Professional Career
1994 - present | Corresponding Member of the Austrian Academy of Sciences |
1991-2002 | Director of Konrad Lorenz Institute for Ethology, Austrian Academy of Sciences, Vienna |
1980-present | Adjunct Professor, Institute of Zoology/ |
1974-1984 | Institute of Limnology of the Austrian Academy of Sciences |
1972-1981 | Lecturer (Animal Behaviour), University of Salzburg |
1973-1974 | Chapman Fellow, Department of Ornithology, Amercian Museum of Natural History, New York |
1972-1973 | Institute of Limnology of the Austrian Academy of Sciences |
1969-1972 | Institute for Comparative Ethology, Austrian Academy of Sciences, Vienna |
Publications
Baldwin, M.W., Winkler, H., Organ, C.L. & Helm, B. (2010): Wing pointedness associated with migratory distance in common-garden and comparative studies of stonechats (Saxicola torquata). Journal of Evolutionary Biology 23: 1050-1063 DOI: 10.1111/j.1420-9101.2010.01975.x [Link 3]
Duda, M., Sattmann, H., Haring, E., Bartel, D., Winkler, H., Harl, J. & Kruckenhauser, L. (2010): Genetic differentiation and shell morphology of Trochulus oreinos (Wagner, 1915) and Trochulus hispidus (Linnaeus, 1785) (Pulmonata: Hygromiidae) in the Northeastern Alps. J. Mollusc. Stud. 77 (1), 33-40. DOI: 10.1093/mollus/eyq037 [Link 4]
Hering, J., Fuchs, E. & Winkler, H. (2010): Neues zum Vorkommen und zur Brutbiologie von Zimtrohrsänger Acrocephalus baeticatus und Teichrohrsänger A. scirpaceus in Libyen. Limicola 24: 117-139.
Raab, R., Kollar, H.P., Winkler, H., Faragó, S., Spakovszky, P., Chavko, J., Maderic, B., Sorpíková, V., Patak, E., Wurm, H., Julius, E., Raab, S. & C. Schütz (2010): Die Bestandsentwicklung der Großtrappe, Otis tarda Linnaeus 1758, von 1900 bis zum Winter 2008/2009. Egretta 51: 74–99.
Winkler, H. & K. Schulze-Hagen (2010): Jürgen Haffer (1932-2010). Vogelwarte 48: 300-301.
Leisler, B. & Winkler, H. (2009): On the evolution of characters associated with migration. Contributions to Natural History 12: 875-892.
Wingelmaier, K., Winkler, H. & Nemeth, E. (2007): Reed bunting (Emberiza schoeniclus) males sing an 'all-clear' signal to their incubating females. Behaviour 144: 195-206.
Kleindorfer, S., Chapman, T. W., Winkler, H. & Sulloway, F. J. (2006): Adaptive divergence in contiguous populations of Darwin's small ground finch (Geospiza fuliginosa). Evol. Ecol. Res. 8: 357-372.
Leisler, B. & Winkler, H. (2006): Interactions between morphological integration and ecological factors. Acta Zoologica Sinica 52: Suppl., 498-501.
Nemeth, E., Dabelsteen, T., Pedersen, S. B. & Winkler, H. (2006): Rainforests as concert halls for birds: Are reverberations improving sound transmission of long song elements? J. Acoust. Soc. Am. 119: 1, 620-626.
Pechacek, P., Michalek, K., Winkler, H. & Blomqvist, D. (2006): Classical polyandry found in the three-toed Woodpecker Picoides tridactylus. Journal of Ornithology 147 (1): 112-114.
Spitzenberger, F., Strelkov, P. P., Winkler, H. & Haring, E. (2006): A preliminary revision of the genus Plecotus (Chiroptera, Vespertilionidae) based on gentic and morphological results. Zool. Scripta 35: 187-230.
Mettke-Hoffmann, C., Wink, M. & Winkler, H. (2005): Exploration of environmental changes relates to lifestyle. Behavioral Ecology 16: 1, 247-254.
Pechacek, P., Michalek, K., Winkler, H. & Blomqvist, D. (2005): Monogamy with exceptions: Social and genetic mating system in a bird species with high paternal investment. Behaviour 142: 8, 1093-1114.
Winkler, H. (2005): Woodpecker coloration. www.ibirding.com/bulletin/articles/V6-wpcolors.pdf [Link 5]
Winkler, H., Kotaka, N., Gamauf, A., Nittinger, F. & Haring, E. (2005): On the phylogenetic position of the Okinawa woodpecker (Sapheopipo noguchii). Journal für Ornithologie 146: 103-110.
Batáry, P., Winkler, H. & Báldi, A. (2004): Experiments with artificial nests on predation in reed habitats. Journal für Ornithologie 145: 59-63.
Kruckenhauser, L., Haring, E., Pinsker, W., Riesling, M. J., Winkler, H., Wink, M. & Gamauf, A. (2004): Genetic vs. morphological differentation of Old World Buzzards (genus Buteo, Accipitridae). Zool. Scripta 33: 197-211.
Schmidt, V., Schaefer, H. M. & Winkler, H. (2004): Conspicuousness, not colour as foraging cue in plant-animal signalling. Oikos 106: 3, 551-557.
Tebbich, S., Taborsky, M., Fessl, B., Dvorak, M. & Winkler, H. (2004): Feeding behavior of four aboreal Darwin 's Finches: Adaptations to spatial and seasonal variability. Condor 106: 95-105.
Winkler, H., Leisler, B. & Bernroider, G. (2004): Ecological constraints on the evolution of avian brains. Journal für Ornithologie 145: 238-244.
Heindl, M. & Winkler, H. (2003): Female canaries (Serinus canaria) associate more with males which contrast strongly against the background. Ethology 109: 259-271.
Heindl, M. & Winkler, H. (2003): Interacting effects of ambient light and plumage color patterns in displaying Wire-tailed Manakins (Aves, Pipridae). Behavioral Ecology and Sociobiology 53: 3, 153-162.
Heindl, M. & Winkler, H. (2003): Vertical lek placement of forest-dwelling manakin species (Aves, Pipridae) is associated with vertical gradients of ambient light. Biol. J. Linn. Soc. 80: 647-658.
Hirschenhauser, K., Winkler, H. & Oliveira, R. F. (2003): Comparative analysis of male androgen responsiveness to social environment in birds: the effects of mating system and paternal incubation. Hormones and Behavior 43: 508-519
Schaefer, H. M., Schmidt, V. & Winkler, H. (2003): Testing the defence trade-off hypothesis: How contents of nutrients and secondary compounds affect fruit removal. Oikos 102: 318-328.
Mettke-Hoffmann, C., Winkler, H. & Leisler, B. (2002): The significance of ecological factors for exploration and neophobia in parrots. Ethology 108: 249-272.
Leisler, B. & Winkler, H. (2001): Morphological convergence in papyrus dwelling passerines. Ostrich Suppl. 15: 24-29
Michalek, K. G. & Winkler, H. (2001): Parental care and parentage in monogamous great spotted woodpeckers (Picoides major) and middle spotted woodpeckers (Picoides medius). Behaviour 138: 1259-1285
Nemeth, E., Winkler, H. & Dabelsteen, T. (2001): Differential degradation of antbird songs in a Neotropical reinforest: Adaptation to perch height? J. Acoust. Soc. Am 110: 6, 3263-327.
Winkler, H. & Preleuthner, M. (2001): Behaviour and ecology of birds in tropical rain forest canopies. Plant Ecology 153: 193-202
Winkler, H. & Kothbauer-Hellmann, R. (2001): The role of search area in the detection of cryptic prey by crested tits and coal tits. Behaviour 138: 873-883
Gamauf, A., Preleuthner, M. & Winkler, H. (1998): Philippine birds of prey: interrelations among habitat, morphology, and behavior. The Auk 115(3): 713-726
Catchpole, C., Leisler, B. & Winkler, H. (1985): Polygyny in the great reed warbler, Acrocephalus arundinaceus : a possible case of deception. Behavioral Ecology and Sociobiology 16 (3): 285-291
Herzig-Straschil, B., Herzig, A. & Winkler, H. (1991): A morpho-metric analysis of the skulls of Xerus inauris and Xerus princeps (Rodentia; Sciuridae). Z. Säugetierkde. 56: 177-187
Hoi, H. & Winkler, H. (1994): Predation on nests: a case of apparent competition. Oecologia 98: 436-440
Kothbauer-Hellmann, R. & Winkler, H. (1997): Prey detection in two tit species Parus ater and P. cristatus. Ethology 103: 339-349
Mbahinzireki, G., Uiblein, F. & Winkler, H. (1991): Microhabitat selection of ostracods in relation to predation and food. Hydrobiologia 222: 115-119
Podolszky, D., Uiblein, F. & Winkler, H. (1995): Visual habitat choice in cyprinid fishes: an experimental approach. Ecology of Freshwater Fish 4: 160-167
Tebbich, S., Taborsky, M. & Winkler, H. (1996): Social manipulation causes cooperation in keas. Animal Behaviour 42: 1-1
Uiblein, F., Eberstaller, J., Plöckl, M. & Winkler, H. (1992): Effects of differential prey mobility on the foraging behaviour of a cyprinid fish, Vimba elongata. Ethology Ecology & Evolution 4: 293-297
Winkler, H. & Orellana, C. P. (1992): The costs of mixed prey for a planktivorous cyprinid, Chalcalburnus chalcoides mento. Environmental Biology of Fishes 35: 85-93
Winkler, H. & Leisler, B. (1992): On the ecomorphology of migrants. Ibis 134 (suppl. 1): 21-28
Winkler, H. & Orellana, C. P. (1992): Functional responses of five cyprinid species to planctonic prey. Environmental Biology of Fishes 33: 53-62
Books and Chapters
Scheifinger, H., Koch, E. & Winkler, H. (2007): Erste Ergebnisse einer Analyse vogelphänologischer Beobachtungen der Zentralanstalt für Meteorologie und Geodynamik 1951-1999 in Österreich. promet 33: 1/2, 52-55.
Winkler, H. & Leisler, B. (2005): To be a migrant - Birds of two worlds. Baltimore, Smithsonian Institution, 79-86
Leisler, B., Forstmeier, W. & Winkler, H. (2004): Kleine Unterschiede im Körperbau - große Effekte. Max-Planck-Gesellschaft Tätigkeitsbericht 2003: 6 pp.
Kurzman, C., Winkler, H., & Steele, B. B. (2003): Feinddruck auf Vogelnester im Waldrandökoton. In: Janauer, G. A. and Hary, N. (Eds.): In: Ökotone Donau-March, Veröffentlichungen des Österreichischen MaB-Programmes. pp: 308, Universitätsverlag Wagner, Innsbruck.
Leisler, B. & Winkler, H. (2003): Morphological consequences of migration in passerines. In: Berthold, P., Gwinner, E., & Sonnenschein, E. (Eds.): Avian migration. pp: 175-186, Springer, Berlin, Heidelberg.
Winkler, Hans & Listabarth, Christian (2003): Surumoni project, Venezuela. In: Basset, Yves, Horlyck, Vibke, & Wright, J. S. (Eds.): Studying forest canopies from above: The international canopy crane network. pp: 126-132, Smithsonian Tropical Research Institute (Panama).
Winkler, H. & Christie, D. (2002): Order Piciformes; Family Picidae. In: Hoyo, J. del, Elliott, A., & Sargatal, J. (Eds.): Handbook of the Birds of the World, pp: 296-555, Lynx Edicions, Barcelona.
Winkler, H. (2002): Sampling birds in the canopy. In: Mitchell, A. W., Secoy, K., & Jackson, T. (Eds.): Global Canopy Handbook, pp: 153-155, Global Canopy Foundation, Oxford.
Winkler, H. & Listabarth, C. (2002): Surumoni canopy crane, Venezuela. In: Mitchell, A. W., Secoy, K., & Jackson, T. (Eds.): Global Canopy Handbook, pp: 80-84, Global Canopy Foundation, Oxford.
Hoi, H., Romero-Pujante, M., & Winkler, H. (2001): The role of tail length for flight performance in the bearded tit (Panurus biarmicus): A tail elongation experiment. In: Hoi, H. (Eds.): The ecology of reed birds, pp: 151-160, Österr. Akademie d. Wissenschaften, Wien.
Winkler, H. (2001): Der Flug der Vögel: von Lorenz bis zur modernen Ökomorphologie. In: Kotrschal, Kurt, Müller, Gerd, & Winkler, Hans (Eds.): Konrad Lorenz und seine verhaltensbiologischen Konzepte aus heutiger Sicht, pp: 195-210, Filander Verlag, Fürth.
Winkler, H. (2001): The ecology of avian acoustical signals. In: Barth, F. G. & Schmid, A. (Eds.): Ecology of sensing, pp: 79-104, Springer-Verlag, Berlin, Heidelberg
Aubrecht, G. & Winkler, H. (2000): Analyse der internationalen Wasservogelzählungen (IWC) in Österreich 1970-1995 - Trends und Bestände. Analysis of the international waterbird cencus (IWC) in Austria 1970-1995 - trends and numbers., pp: 175, Ser.: Morawetz, W. and Winkler, H. (Ser. Eds.): Biosystematics and ecology series No 13, Österr. Akad. d. Wissenschaften, Wien.
Hille, S. & Winkler, H. (2000): Ecomorphology of island populations of the Kestrel (Falco tinnunculus) on Cape Verde. In: Chancellor, R. D. (Eds.): Raptors at risk, pp: 729-736, WWWGBP/Hancock House, Blaine, WA.
Preleuthner, M. & Winkler, H. (2000): Neue Forschungsmethoden im Regenwald - Mit dem Kran in die Kronen.: 23. Int. Symp. Mensch-Tier-Umwelt 1999, pp: 34-37.
Winkler, H. & Zwicker, E. (2000): Beziehungen zwischen Vogelgemeinschaften und Vegetationsstrukturen und deren Beeinflussung durch Forstwirtschaft und Kraftwerk. In: Parz-Gollner, R & Herzig, A. (Eds.): Ökosystemstudie Donaustau Altenwörth, pp: 214-231.
Zwicker, E. & Winkler, H. (2000): Die Auswirkung wasserbaulicher Maßnahmen auf die Vogelwelt der Augewässer. In: Parz-Gollner, R & Herzig, A. (Eds.): Ökosystemstudie Donaustau Altenwörth, pp: 115-124.
Leisler, B. & Winkler, H. (1985): Ecomorphology. In: Johnston, R. F. (Eds.), pp: 155-186, Plenum Press, New York, London.
Newton, S. F., Newton, A. V., Winkler, H., & Orellana, C. P. (1994): The avifauna of Wadi Turabah: Status, distribution and habitat associations. In: Büttiker, W. & Krupp, F. (Eds.): Fauna of Saudi Arabia. Vol. 14, pp: 442-454, Pro Entomologia, Basel.
Winkler, H. & Leisler, B. (1985): Morphological aspects of habitat selection in birds. In: Cody, M. L. (Eds.): Habitat selection in birds, pp: 415-434.
Winkler, H. (1990): Probleme des terrestrischen Biotopschutzes im Neusiedler See-Gebiet. In: Arbeitsgemeinschaft Gesamtkonzept, Neusiedler See (Eds.): Tagungsband Int. Symp. 'Schutz und Entwicklung großer mitteleuropäischer Binnenseenlandschaften. Bodensee - Neusiedler See - Balaton', 24.-27. Apr. 1990, pp: 115-124, Pamhagen im Seewinkel (Burgenland).
Winkler, H., Berthold, P., & Leisler, B. (1994): Monitoring of bird populations in the Lake Neusiedl area. In: Aubrecht, G., Dick, G., & Prentice, C. (Eds.): Monitoring of Ecological Change in Wetlands of Middle Europe. Int. Workshop, Linz, Austria, 1993, pp: 29-36, IWRB Publ. No 30, Slimbridge, UK.
Winkler, H. (1994): Tierisches Verhalten - ein Motor der Evolution. In: Wieser, W. (Eds.): Die Evolution der Evolutionstheorie, pp: 194-220, Spektrum Akad. Verl., Heidelberg.
Winkler, H., Christie, D. A., & Nurney, D. (1995): Woodpeckers. Guide to the woodpeckers, piculets and wrynecks of the world., pp: 406, Pica Press., Sussex.
Abstracts
Hille, S., Blomqvist, D., Nesje, M. & Winkler, H. (2002): Morphological, behavioural, and genetic variation in the Cape Verde Kestrels (Falco tinnunculus). Raptors in the new millenium Proc. of the world conference on birds of prey & owls., In: Yosef, R., Miller, M. L., and Pepler, D., p. 220, Raptor Research Foundation, Eilat.
Heindl, M. & Winkler, H. (2000): Interaction of ambient light and plumage color in the Wire-tailed Manakin (Pipra filicauda , Pipridae, Aves). Advances in Ethology 35 (Suppl. 38): p. 20
Hille, S. & Winkler, H. (2000): Adaptive behavior of island populations of the Kestrel (Falco tinnunculus) to heterogenous environments. Advances in Ethology 35 (Suppl.): p. 39
Winkler, H. & Leisler, B. (1999): Exploration and curiosity in birds: Functions and mechanisms. Acta 22 Congr. Int. Ornithol., In: Adams, N. J. and Slotow, R. H., p. 915-932, BirdLife South Africa, Johannesburg, Durban.
Gamauf, A., Preleuthner, M. & Winkler, H. (1998): Mimicry in SE-Asian Birds of Prey: Barred Honey Buzzards (Pernis celebensis) vs. Hawk-Eagles (Spizaetus spp.). 5th World Conference on Birds of Prey and Owls, Midrand, Johannesburg, Südafrika.
Preleuthner, M., Grubbauer, P. & Winkler, H. (1998): Are canopy birds ethologically and ecologically distinct? Forest Canopies 1998: Global Perspectives. Sarasota, Florida.
Preleuthner, M., Grubbauer, P. & Winkler, H. (1998): Ist die Avifauna des Kronendachs ethologisch und ökologisch distinkt? 11. Jahrestagung der Deutschen Ges. f. Tropenökologie. Bd 12: In: Dalitz, H., Haverkamp, M., Homeier, J., and Breckle, S. W.: Bielefelder Ökologische Beiträge, Bielefeld, Deutschland.
Leisler, B. & Winkler, H. (1994): Morphological aspects of habitat selection. Journal für Ornithologie 135: p. 482-483
Winkler, H. (1988): An examination of concepts and methods in ecomorphology. Acta XIX Congressus Int. Ornithologici II, 2246-2253
Winkler, H. (1990): Verhalten als Mittler zwischen Morphologie und Habitat. Proc. Int. 100. DO-G Meeting, Current Topics Avian Biol.; Bonn, p. 345-350
Winkler, H. & Leisler, B. (1994): Ecomorphological patterns of adaptation and convergence in forest birds. Journal für Ornithologie 135: p. 472-473
Educational, Popular and Other
Scheifinger, H., Koch, E. & Winkler, H. (2005): Results of a first look into the Austrian animal phenological records. Meteorolog. ZS 14: 2, 1-7.
Heindl, M. & Winkler, H. (2003): Interaction of ambient light, lek placement, display behavior, and color signals in two Pipra species. 23rd International Ornithological Congress, Beijing
Leisler, B. & Winkler, H. (2003): Morphological convergence in papyrus dwelling passerines. Bonner Zoologische Beiträge 51: 119
Michalek, K., Auer, J. A., Großberger, H., Schmalzer, A. & Winkler, H. (2001): Die Einflüsse von Lebensraum, Witterung und Waldbewirtschaftung auf die Brutdichte von Bunt- und Mittelspecht (Picoides major und P. medius) im Wienerwald. Influence of habitat, wether and forest managment on breeding density of Great Spotted Woodpeckers and Middle Spotted Woodpeckers (Picoides major und P. medius) in the Vienna Woods ("Wienerwald"). Abh. Ber. Mus. Heineanum 5: Sonderheft, 31-58
Preleuthner, M. & Winkler, H. (2000): Projekt Surumoni Österreichische Kronendach-Forschung im größten Regenwaldgebiet der Erde. Schönbrunner Tiergartenjournal 3/2000: 1-8
Anhuf, D. & Winkler, H. (1999): Geographical and ecological settings of the Surumoni-Crane Project (Upper Orinoco, Estado Amazonas, Venezuela). Anzeiger der Mathematisch-Naturwissenschaftlichen Klasse Abteilung I 135: 3-23
Michalek, K., Auer, J. A. & Winkler, H. (1999): Natal dispersal and returning of former nestlings in monogamous Great Spotted Woodpeckers (Picoides major) and Middle Spotted Woodpeckers (Picoides medius).Dispersion der Jungvögel und Rückkehr der Nestlinge beim Buntspecht (Picoides major) und beim Mittelspecht (Picoides medius). Tichodroma 12: Suppl. 1, 122-133
Winkler, H. & Preleuthner, M. (1999): The ecomorphology of Neotropical frugivores. Acta Ornothologica 34: 2, 141-148
Winkler, H. (1998): Die Rolle des Verhaltens für die Artbildung der Vögel. Zool. Abh. Staatl. Mus. f. Tierkde. Dresden 50/Suppl:2: 21-33
Bairlein, A., Leisler, B. & Winkler, H. (1986): Morphologische Aspekte der Habitatwahl von Zugvögeln in einem SW-deutschen Rastgebiet. Journal für Ornithologie 127(4): 463-473
Berthold, P., Fliege, G., Querner, U. & Winkler, H. (1986): Die Bestandsentwicklung von Kleinvögeln in Mitteleuropa Analyse von Fangzahlen. Journal für Ornithologie 127 (4): 397-437
Berthold, P., Querner, U. & Winkler, H. (1988): Vogelschutz: 100 Jahre lang bis in die "roten Zahlen" - ein neues Konzept ist unerläßlich. Natur u. Landschaft 63 (1): 5-8
Herzig, A. & Winkler, H. (1985): Der Einfluß der Temperatur auf die embryonale Entwicklung der Cypriniden. Öst. Fischerei 38: 182-196
Hoi, H. & Winkler, H. (1988): Feinddruck auf Schilfbrüter: Eine experimentelle Untersuchung. Journal für Ornithologie 129 (4): 439-447
Hoi, H. & Winkler, H. (1991): Feinddruck auf Vogelnester: Effekte der Habitatkomplexität. Journal für Ornithologie 132 (2): 215-217
Kothbauer-Hellmann, R. & Winkler, H. (1991): Aufenthaltsdauern explorierender Meisen: Habitat- und Artspezifität. Journal für Ornithologie 132 (3): 329-331
Kurzman, C. & Winkler, H. (1994): Predation pressure on nests in woodland ecotones: effect of edge distance. Journal für Ornithologie 135 (Sonderheft): p. 198
Leisler, B., Ley, H. W. & Winkler, H. (1987): Die Rolle von Lokomotion und Morphologie in der Habitatwahl der Rohrsänger (Acrocephalus). Journal für Ornithologie 128 (1): 115-117
Leisler, B., Ley, H. W. & Winkler, H. (1989): Habitat, behaviour and morphology of Acrocephalus warblers: an integrated analysis. Ornis Scand. 20: 181-186
Leisler, B. & Winkler, H. (1991): Ergebnisse und Konzepte ökomorphologischer Untersuchungen an Vögeln. Journal für Ornithologie 132 (4): 373-425
Leisler, B., Winkler, H. & Siebenrock, K. H. (1997): Ökomorphologische Untersuchungen am Beispiel der Webervögel (Ploceidae) und Eisvögel (Alcedinidae). Mitt. Zool. Mus. Berl. 73 Suppl: Ann. Orn. 21: 17-43
Michalek, K. & Winkler, H. (1997): Der Buntspecht - Vogel des Jahres 1997 - Hacken und Klettern - ein Leben am Baum. Falke 44: 4-9
Ritzmeier, M. & Winkler, H. (1994): Effects of hunger and protein on exploration. Journal für Ornithologie 135 (Sonderheft): p. 150
Schulze-Hagen, K., Leisler, B. & Winkler, H. (1992): Teich- und Sumpfrohrsänger in gemeinsamem Habitat: Brutzeit und Bruterfolg. Vogelwelt 113: 89-98
Schulze-Hagen, K., Leisler, B. & Winkler, H. (1996): Breeding success and reproductive strategies of two Acrocephalus warblers. Journal für Ornithologie 137: 181-192
Symens, P., Newton, S. F., Winkler, H. & Stagg, A. J. (1994): Mountain nightjar Caprimulgus poliocephalus in Arabia: identification, status, and distribution. Sandgrouse 14: 81-92
Taborsky, M. & Lindauer, M. (1993): [Bookreview] Auf den Spuren des Uneigennützigen (München: Artemis & Winkler 1991). Universitas 48 (1): 87-88
Uiblein, F. & Winkler, H. (1994): Der Nahrungserwerb der Mondsee Zährte. Sitzungsber. Österr. Akad. Wiss.; Math. -nat. Kl.; Abt. I 196: 311-322
Uiblein, F. & Winkler, H. (1994): Morphological variability among Vimba in Austrian waters: quantitative examination of a taxonomic and a functional hypothesis (Pisces: Cyprinidae). Senckenbergiana biologica 73 (1/2): 57-65
Uiblein, F., Rydlo, M., Kainz, E. & Winkler, H. (1996): Welche Funktion hat die Laichfärbung der Zährte (Vimba sp.; Cyprinidae)? Öst. Fischerei 49: 220-228
Winkler, H. & Moreno, T. (1984): Die funktionelle Reaktion planktonfressender Fische. Sitzungsber. Österr. Akad. Wiss.; Math. -nat. Kl.; Abt. I 193: p. 23-31
Winkler, H. (1985): Eine Methode zur Bestimmung von Aktivitätsmustern aus Felddaten. Vogelwarte 33 (2): 156-161
Winkler, H. (1988): Paarungssysteme: Neuere Ergebnisse der Verhaltensforschung. Wiss. Nachrichten (B. M. f. Unterr.; Kunst u. Sport) 77 (4): 11-13
Winkler, H. (1989): Prinzipien der Lebensraumgestaltung. Der Aufbau - Perspektiven l989 62-64
Winkler, H. (1991): Verhaltensökologie und Biotopschutz: von der Theorie zur Praxis. Seevögel 12: (Sonderh.1), 115-118
Winkler, H. (1992): A study of the Arabian Woodpecker (Picoides dorae) an Arabian endemic (6 March to 5 Apr. 1992). Research Report 1-9
Winkler, H., Christie, D. A. & Nurney, D. (1994): The colourful world of woodpeckers: an Oriental perspective. OBC (Oriental Bird Club) Bull. 19: May, Spec. iss., 30-33
Winkler, H., Newton, A. V. & Newton, S. F. (1996): On the ecology and behaviour of the Arabian Woodpecker Picoides dorae. Zool. Middle East 12: 33-45
Zwicker, E. & Winkler, H. (1989): Einige Auswirkungen wasserbaulicher Maßnahmen auf die Vogelwelt der Ufersäume von Augewässern. Öst. Wasserwirtschaft 41: 215